Weed control pushes populations toward the crop niche. Weed fitness pulls them back. Explore the balance.
Control selection removes weeds probabilistically. Further from the crop = higher kill probability, but chance means some distant weeds survive.
Fitness cost penalises reproduction for weeds that deviate from their own optimum. At zero, convergence is inevitable. At high values, populations equilibrate partway.
IWM rotates the selection zone each generation. No single trait combination stays safe, disrupting directional convergence.